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Melinda Clarke, 'The O.C.'
Then: She slept with the kid who took her daughter's virginity, married her best friend's father, then almost poisoned him (but he had a heart attack before she got the chance). And yet -- everybody really freaking loved Julie Cooper. So much.
Now: Clarke showed us how Julie would've ended up if she'd stayed in Riverside with a guest-starring arc on "The Vampire Diaries" as Matt's deadbeat mom. This season, she's the HBIC of Division on "Nikita," which airs Fridays at 9 p.m. ET on The CW.